When feeding my rabbits branches or bark chews in the winter, I do feed some spruce and pine in moderation, while being mindful of when I read briefly about pine induced abortion in
cattle. I admittedly didn't read deep into the studies regarding the specific species of evergreen or whether it can occur in other animals. I don't breed my rabbits in the winter months anyway and have never noticed any side effects from feeding spruce boughs. By the time I start breeding in springtime (Maine z5), other trees are budding.
According to
Bunnymeadow spruce is safe.