I've focused this year on the 3/4 domestics that seemed so promising in Joseph's garden last year. The best of them are going into the fenced garden. Have some planted and some more seed planted (G3) and garden space held for those. Waiting for germination!
The seed I grew of 3/4 wilds went into the general population as a frost test, a regular plant out, and a direct seeding. I expect it to segregate into edibility at a higher rate than before but not as high as the plants that were best from 2019 in Josephs larger growout.
Then I also in the general population have some pure habrochaites of a new strain, pure peruvianum, and pure arcanum. I have what I believe to be high percentage penellii and high percentage habrochaites from wild maternal cytoplasm as well. Also a three way cross Joseph sent that has domestic maternal cytoplasm.
Then I have a mini population with lots of penellii and one peruvianum. I hope that it will result in a good amount of presumptive penellii x peruvianum seed with peruvianum as the mother for 2021.
I haven't grown any more chilense this year which is probably the most promising germ plasm for crossing with peruvianum I have. It's also the tomato hardest for me to cultivate based on my 2019 results which was plants that died young before blooming.
I also have all thoughts about making embryo rescue peruvianum on hold till a time when I have more time.
I'm very curious about the crossed accessions peruvianum x domestic Andrew mentions. Particularly in terms of generation?
I intend to plant an exserted domestic right next to the Arcanum but not sure if I'll get it done or be able to grow the resulting seed enmasse if I do. A controlled Arcanum x domestic cross should be possible and would result in another interesting line of its own. However is not reportedly a good bridge after all but rather something interesting in its own right.
Roads not travelled this year include not growing many of the exciting penellii germ plasm from last year yet. Though there is penellii in the 3/4 domestic lines and Joseph thought it produced some of the best 2019 flavors. I counted about nine plants with penellii leaf surface dots in the best flavor population. I meant to sort them but didn't. Small possibility there that crossing behavior may be directional based on something Andrew and Joseph were pointing out the other day. Penellii can in theory pollinate domestic, habrochaites, and peruvianum. However perhaps not the reverse. If the hybrids behave at least partially in the same way we should expect penellii hybrids to pollinate: penellii hybrids, perhaps back pollinate pure penellii (we think we have such a population, domestics, habrochaites and hab hybrids, and maybe peruvianum. However habrochaites hybrids probably cannot pollinate penellii hybrids. Though this thought may be taking that thinking to far. It's also possible that complex penellii x hab x domestic hybrids can pollinate in both directions depending on which genes are present and from whom. Though I do suspect that in our multiple wild species promiscous lines there is probably some odd crossing behavior that may be undetectable. If we add more species it may further complicate that within a general multi species hybrid zone population. Which makes me wonder if we need multiple populations with 150 feet of separation between. Even in the 3/4 domestic population wonder if we should separate out the penellii influenced portion and grow it by itself? Also makes me wonder if we could ultimately have multiple promiscuous populations derived from different parental stock that still behave like separate species. I.e. penellii based, habrochaites based, peruvianum based, and arcanum based? As opposed to a universal hybrid zone in which you might end up with plants with small introgressions from all species like modern domestic tomatoes but uncertain pollen flow behaviors.